Editorial Board

EDITOR IN CHIEF (2016)

Harold L. Drake

University of Bayreuth

EDITORS

Thomas E. Besser, Editor (2020)

Washington State University

Johanna Björkroth, Editor (2018)

University of Helsinki

Axel A. Brakhage, Editor (2016)

HKI and University of Jena

Daniel Cullen, Editor (2017)

USDA Forest Products Laboratory

Charles M. Dozois, Editor (2020)

INRS—Institut Armand-Frappier

Edward G. Dudley, Editor (2020)

Pennsylvania State University

Christopher A. Elkins, Editor (2018)

U.S. FDA Center for Food Safety and Applied Nutrition

Marie A. Elliot, Editor (2019)

McMaster University

Heidi Goodrich-Blair, Editor (2017)

University of Wisconsin—Madison

Robert M. Kelly, Editor (2017)

North Carolina State University

Maia Kivisaar, Editor (2017)

Tartu University

Joel E. Kostka, Editor (2016)

Georgia Institute of Technology

Shuang-Jiang Liu, Editor (2016)

Chinese Academy of Sciences

Frank E. Löffler, Editor (2018)

University of Tennessee

Volker Müller, Editor (2019)

Goethe University Frankfurt am Main

Hideaki Nojiri, Editor (2017)

The University of Tokyo

Rebecca E. Parales, Editor (2016)

University of California—Davis

M. Julia Pettinari, Editor (2016)

University of Buenos Aires

Donald W. Schaffner, Editor (2020)

Rutgers, The State University of New Jersey

Patrick D. Schloss, Editor and Cover Editor (2019)

University of Michigan

Janet L. Schottel, Editor (2018)

University of Minnesota

Alfred M. Spormann, Editor (2016)

Stanford University

Alfons J. M. Stams, Editor (2020)

Wageningen University

Claire Vieille, Editor (2020)

Michigan State University

Gerrit Voordouw, Editor (2017)

University of Calgary

Thomas E. Shenk, Chairman, Journals Board

Barbara M. Goldman, Director, Journals

Barbara S. Slinker, Production Editor

Stephanie L. D'Agostino, Assistant Production Editor

Michael E. Lerman, Assistant Production Editor

EDITORIAL BOARD

Adly Abd-Alla (2015)

Lorenz Adrian (2017)

Warish Ahmed (2016)

Birgit E. Alber (2016)

Gladys Alexandre (2015)

Kevin J. Allen (2015)

Michael S. Allen (2017)

Eric Altermann (2015)

Gary L. Andersen (2016)

Matthew J. Arduino (2016)

Arthur I. Aronson (2015)

Fred O. Asiegbu (2016)

Haruyuki Atomi (2016)

Shota Atsumi (2015)

Beate Averhoff (2016)

Abdu F. Azad (2016)

M. Andrea Azcarate-Peril (2015)

Reha O. Azizoglu (2015)

Charles W. Bacon (2015)

Petr Baldrian (2016)

Jeffrey Banas (2015)

Ravi D. Barabote (2015)

Jeri D. Barak (2016)

Rodolphe Barrangou (2017)

Florian F. Bauer (2015)

Jorge A. Benitez (2016)

Teresa M. Bergholz (2015)

Stefan Bertilsson (2015)

Arvind A. Bhagwat (2015)

Valeria Bianciotto (2015)

Kelly A. Bidle (2016)

Linda F. Bisson (2016)

Christopher Blackwood (2016)

Melanie Blokesch (2016)

Paul L. Bodelier (2015)

Eckhard Boles (2015)

Matthias Boll (2015)

Daniel R. Bond (2015)

Albert Bosch (2015)

Peter Bottomley (2015)

Kostas Bourtzis (2015)

Frederick Breidt (2017)

Peter A. Bron (2017)

Harald Bruessow (2016)

Harry Brumer (2017)

Alison Buchan (2017)

Robert L. Buchanan (2015)

Wolfgang Buckel (2016)

Daniel H. Buckley (2017)

William Burkhardt (2015)

Helen Y. Buse (2016)

Stefan G. Buzoianu (2017)

Douglas R. Call (2016)

Amy Callaghan (2015)

Terri A. Camesano (2015)

Barbara J. Campbell (2016)

Isaac K. O. Cann (2015)

Jennifer L. Cannon (2017)

Yiping Cao (2017)

Michelle Q. Carter (2015)

Rick Cavicchioli (2016)

Shane M. Ceraul (2015)

Woo-Suk Chang (2015)

Jinru Chen (2017)

Shicheng Chen (2017)

Wilfred Chen (2015)

Yi Chen (2017)

Yuhuan Chen (2015)

Michael L. Chikindas (2016)

Ludmila Chistoserdova (2015)

Kuang-Ren Chung (2017)

Clifford G. Clark (2017)

John D. Coates (2016)

Luca Cocolin (2016)

Doug Cossar (2015)

Paul D. Cotter (2016)

Don A. Cowan (2015)

Alison M. Cupples (2016)

Holger Daims (2015)

Rolf Daniel (2015)

Steven L. Daniel (2017)

Michelle Danyluk (2016)

Jennifer M. DeBruyn (2017)

Svetlana Dedysh (2015)

Zixin Deng (2016)

Angelo DePaola, Jr. (2015)

Mickaël Desvaux (2015)

Thomas J. DiChristina (2015)

Francisco Diez-Gonzalez (2016)

Lubbert Dijkhuizen (2016)

Jayna L. Ditty (2016)

Jan Dolfing (2015)

James S. G. Dooley (2015)

Angela E. Douglas (2016)

Irina S. Druzhinina (2015)

Doris d'Souza (2017)

Alain C. Dufour (2016)

Jonathan Dworkin (2017)

Daniel C. Eastwood (2017)

Elizabeth A. Edwards (2016)

Henry L. Ehrlich (2015)

Patrick Y. Eichenberger (2015)

Mostafa Elshahed (2015)

David Emerson (2015)

Slava Epstein (2016)

Tobias J. Erb (2017)

Danilo Ercolini (2015)

Joseph O. Falkinham (2017)

Seamus Fanning (2015)

Babu Z. Fathepure (2016)

J. Christopher Fenno (2016)

Martin Filion (2015)

Steven E. Finkel (2016)

Steven L. Foley (2017)

Philipp Franken (2015)

W. Florian Fricke (2017)

Michael W. Friedrich (2015)

Cormac Gahan (2015)

Victor P. J. Gannon (2016)

Michael G. Gänzle (2016)

Beilei Ge (2016)

Elke Genersch (2015)

Johannes Gescher (2015)

Lutz Geue (2016)

Jennifer B. Glass (2017)

Emanuel Goldman (2016)

Mark Gomelsky (2017)

Ramon Gonzalez (2016)

Lawrence Goodridge (2015)

Lisa A. Gorski (2015)

David E. Graham (2017)

Jeffrey Gralnick (2015)

Stefan J. Green (2017)

Leslie A. Gregg-Jolly (2017)

Richard L. Gregory (2015)

Yitzhak Hadar (2016)

Jan O. Haerter (2015)

Thomas E. Hanson (2015)

Eiji Haramoto (2015)

Hauke Harms (2016)

Steven D. Harris (2017)

Mark E. Hart (2017)

Valerie J. Harwood (2015)

Annele I. Hatakka (2016)

Paul B. Hatzinger (2015)

Qiang He (2016)

Zhili He (2016)

Brian P. Hedlund (2017)

Henry S. Heine (2015)

Niels B. Hendriksen (2015)

Laura Hendrix (2015)

Ute Hentschel (2015)

Salvador Herrero (2015)

Paul S. Hoffman (2015)

Dirk Hoffmeister (2015)

Andrew J. Holmes (2016)

Allen L. Honeyman (2015)

Dallas G. Hoover (2015)

David J. Hopper (2016)

Marcus A. Horn (2017)

John A. Hudson (2017)

Jeroen Hugenholtz (2015)

Hor-Gil Hur (2015)

Michael R. Hyman (2016)

Michael F. Hynes (2016)

Kiyohiko Igarashi (2016)

Michele M. Igo (2015)

Masaharu Ishii (2017)

Masahiro Ito (2017)

Paul J. Jackson (2015)

Mohamed A. Jamal (2015)

Edward L. Jarroll (2015)

Dieter Jendrossek (2015)

Ian Jenson (2017)

Zhongjun Jia (2017)

Nianzhi Jiao (2017)

Zackary I. Johnson (2015)

Yves Jouanneau (2015)

Juan L. Jurat-Fuentes (2015)

Boran Kartal (2017)

Matthias Kästner (2016)

James E. Keen (2016)

John J. Kelly (2015)

William Kenealy (2015)

Eungbin Kim (2015)

James E. Kirby (2016)

Arnulf Kletzin (2016)

Leon D. Kluskens (2016)

Kalmia E. Kniel (2017)

Steffen Kolb (2017)

Konstantinos T. Konstantinidis (2015)

Shige Koseki (2015)

Michael Y. Kosoy (2017)

Balasubrahmanyam Kottapalli (2017)

George Kowalchuk (2015)

Sven Krappmann (2015)

Lee R. Krumholz (2015)

Indira T. Kudva (2016)

Ursula Kües (2016)

Timothy M. LaPara (2015)

Gisèle LaPointe (2017)

Robert A. LaRossa (2015)

Luis F. Larrondo (2017)

Peter C. K. Lau (2015)

Jared R. Leadbetter (2015)

Jeffrey T. LeJeune (2015)

Paulo C. Lemos (2015)

Steven Lindow (2016)

John T. Lisle (2015)

Wenjun Liu (2016)

Xueduan Liu (2016)

Karen Lloyd (2015)

Martin J. Loessner (2015)

John C. Londesborough (2015)

Alexander Loy (2017)

Yahai Lu (2016)

Tillmann Lueders (2015)

Yanhe Ma (2016)

Sandra Macfarlane (2017)

Robert L. Mach (2015)

Eugene L. Madsen (2015)

Jacques Mahillon (2017)

Mark J. Mandel (2017)

Maria L. Marco (2015)

Helene Marquis (2015)

Matthew J. Marshall (2016)

Stephen Martin (2015)

Alfredo Martinez (2016)

Eiji Masai (2017)

Emma Master (2015)

Timothy E. Mattes (2017)

Ann G. Matthysse (2015)

Andrew J. McBain (2015)

Timothy R. McDermott (2015)

Michael J. McInerney (2016)

James McKinlay (2015)

Robert J. McLean (2015)

Brian B. McSpadden-Gardener (2016)

Rainer U. Meckenstock (2015)

Richard J. Meinersmann (2015)

Aubrey F. Mendonca (2015)

John S. Meschke (2017)

Chris H. Miller (2015)

Patricia D. Millner (2015)

Debra L. Milton (2016)

Kiwamu Minamisawa (2015)

Kentaro Miyazaki (2017)

Corrie S. Moreau (2017)

Rodney A. Moxley (2016)

Jochen A. Mueller (2015)

Yuji Nagata (2015)

Cindy H. Nakatsu (2016)

Brett A. Neilan (2017)

Nancy Nichols (2015)

Brendan A. Niemira (2015)

Pablo I. Nikel (2016)

Nobuhiko Nomura (2017)

R. Sean Norman (2015)

Julie B. Olson (2015)

Huub Op den Camp (2015)

Ronald S. Oremland (2015)

Ivan J. Oresnik (2017)

Konstantinos Papadimitriou (2016)

Hee-Deung Park (2017)

Ilenys M. Perez-Diaz (2015)

Gregg S. Pettis (2017)

Jennifer Pett-Ridge (2015)

Laurent Philippot (2017)

Allen T. Phillips (2015)

Carmen Pin (2015)

Antonio G. Pisabarro (2017)

Mircea Podar (2017)

David L. Popham (2015)

Rolf Prade (2015)

M. Auxiliadora Prieto (2015)

Qingsheng Qi (2017)

Cheryl L. Quinn (2015)

Ralf Rabus (2017)

Mark Radosevich (2017)

Andreja Rajkovic (2015)

Gemma Reguera (2015)

Bernd H. Rehm (2015)

Gregor Reid (2015)

Christian Riedel (2015)

Kirsti M. Ritalahti (2015)

Daniel D. Rockey (2016)

Jorge L. M. Rodrigues (2016)

David Rodrígues-Lázaro (2015)

Wilfred F. M. Röling (2016)

Margaret F. Romine (2017)

Thomas Ross (2016)

Michael J. Sadowsky (2016)

Daâd A. Saffarini (2017)

Maria Sandkvist (2015)

Robert A. Sanford (2015)

Anderson S. Sant'Ana (2015)

Sima Sariaslani (2015)

Brian D. Sauders (2016)

Gary Sawers (2017)

Frank A. Scannapieco (2017)

Birgit E. Scharf (2017)

Bernhard Schink (2015)

John W. Schmidt (2016)

Monika Schmoll (2017)

Hildgund Schrempf (2017)

Dirk Schüler (2015)

Wolfgang H. Schwarz (2016)

Lynne Sehulster (2016)

Bernhard Seiboth (2017)

Verena Seidl (2015)

Jeremy D. Semrau (2017)

Peter Setlow (2015)

Joshua D. Shrout (2015)

Mark W. Silby (2015)

Mitchell H. Singer (2017)

Steven W. Singer (2015)

Brajesh K. Singh (2016)

David R. Singleton (2015)

Matthias Sipiczki (2016)

Panos N. Skandamis (2016)

Evan Skowronski (2015)

J. Leif Smith (2016)

Helena Solo-Gabriele (2015)

Greg A. Somerville (2017)

Andrew J. Spiers (2016)

Stephen Spiro (2016)

Eric V. Stabb (2016)

Lisa Y. Stein (2016)

Alexander Steinbüchel (2017)

Ivo Steinmetz (2016)

Roger Stephan (2017)

Ann M. Stevens (2016)

Brian Stevenson (2016)

George C. Stewart (2015)

John F. Stolz (2016)

Wolfgang R. Streit (2015)

Joseph M. Sturino (2016)

Garret Suen (2016)

Matt D. Sweeney (2016)

Ben D. Tall (2017)

Ralph S. Tanner (2017)

Gerald W. Tannock (2017)

Sam R. Telford (2015)

Andreas P. Teske (2015)

Michael G. Thomas (2015)

Kai M. Thormann (2015)

Thomas Thurnheer (2015)

Nicolas Toro Garcia (2017)

Effie Tsakalidou (2016)

Masataka Tsuda (2015)

Hidetoshi Urakawa (2017)

Wouter R. van der Star (2017)

Tina K. Van Dyk (2015)

Jan D. van Elsas (2016)

Michael L. Vasil (2016)

Marco Ventura (2015)

Dmitriy V. Volokhov (2015)

Julia A. Vorholt (2015)

Stéphane Vuilleumier (2015)

S. Aljoscha Wahl (2015)

Judy D. Wall (2015)

R. John Wallace (2016)

Todd J. Ward (2016)

Keith Warriner (2017)

J. Scott Weese (2015)

Helge Weingart (2015)

Alan Welman (2015)

Tingyi Wen (2015)

Volker F. Wendisch (2015)

Giovanni Widmer (2016)

Steven W. Wilhelm (2015)

Kurt E. Williamson (2017)

David B. Wilson (2016)

Charles F. Wimpee (2017)

Regina-Michaela Wittich (2017)

Gideon M. Wolfaardt (2016)

Randy W. Worobo (2016)

Qinglong Wu (2015)

Xiao-Lei Wu (2016)

Chuanwu Xi (2017)

Hua Xiang (2015)

Lihua Xiao (2015)

Luying Xun (2015)

S. Steve Yan (2016)

X. Frank Yang (2015)

Yunfeng Yang (2015)

Süleyman Yildirim (2017)

Daniel R. Zeigler (2015)

Jill H. Zeilstra-Ryalls (2017)

Karsten Zengler (2017)

Chuanlun Zhang (2015)

Qijing Zhang (2017)

Wei Zhang (2015)

Yu-Zhong Zhang (2017)

Ning-Yi Zhou (2015)

Erik R. Zinser (2016)

Erwin G. Zoetendal (2015)

Angeles Zorreguieta (2016)

Sergey B. Zotchev (2015)

Peter A. Zuber (2017)

Applied and Environmental Microbiology (ISSN 0099-2240), a publication of the American Society for Microbiology (ASM), 1752 N St., N.W., Washington, DC 20036-2904, is devoted to the advancement and dissemination of applied knowledge as well as ecological knowledge, both applied and fundamental, concerning microorganisms. Instructions to Authors can be found at http://journalitas.asm.org/t/49536 and are updated throughout the year. Applied and Environmental Microbiology is published twice monthly, one volume per year. Nonmember print subscription prices (per year) are: $1,984, U.S.; $2,103, Canada (plus GST/HST where applicable); $2,136, Europe; $2,189, Latin America; $2,195, rest of world. Member print subscription prices (per year) are: $370, U.S.; $464, Canada (plus GST/HST where applicable); $479, Europe; $517, Latin America; $522, rest of world. Single copies are: $100, nonmember; $50, member (Canadians add GST/HST where applicable). For prices of electronic versions or for correspondence relating to subscriptions, defective copies, missing issues, and availability of back issues, contact the Subscriptions Unit, ASM (asmjournals{at}subscriptionoffice.com). Correspondence relating to reprint orders should be directed to ASM Author Services (ASM.AuthorServicesSupport{at}cenveo.com), and correspondence relating to disposition of submitted manuscripts, proofs, and general editorial matters should be directed to the Journals Department, American Society for Microbiology, 1752 N St., N.W., Washington, DC 20036-2904. Phone: (202) 737-3600 (see http://journals.asm.org/site/misc/staffcontacts.xhtml).

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