Editorial Board

EDITOR IN CHIEF (2021)

Harold L. Drake

University of Bayreuth

EDITORS

Haruyuki Atomi, Editor (2021)

Kyoto University

Johanna Björkroth, Editor (2018)

University of Helsinki

Daniel Cullen, Editor (2017)

USDA Forest Products Laboratory

Charles M. Dozois, Editor (2020)

INRS—Institut Armand-Frappier

Edward G. Dudley, Editor (2020)

Pennsylvania State University

Christopher A. Elkins, Editor (2018)

U.S. FDA Center for Food Safety and Applied Nutrition

Marie A. Elliot, Editor (2019)

McMaster University

Robert M. Kelly, Editor (2017)

North Carolina State University

Maia Kivisaar, Editor (2017)

Tartu University

Joel E. Kostka, Editor (2021)

Georgia Institute of Technology

Shuang-Jiang Liu, Editor (2021)

Chinese Academy of Sciences

Frank E. Löffler, Editor (2018)

University of Tennessee

Emma Master, Editor (2021)

University of Toronto

Andrew J. McBain, Editor (2021)

University of Manchester

Volker Müller, Editor (2019)

Goethe University Frankfurt am Main

Hideaki Nojiri, Editor (2017)

The University of Tokyo

Rebecca E. Parales, Editor (2021)

University of California—Davis

M. Julia Pettinari, Editor (2021)

University of Buenos Aires

Donald W. Schaffner, Editor (2020)

Rutgers, The State University of New Jersey

Patrick D. Schloss, Editor and Cover Editor (2019)

University of Michigan

Janet L. Schottel, Editor (2018)

University of Minnesota

Eric V. Stabb, Editor (2021)

University of Georgia

Alfons J. M. Stams, Editor (2020)

Wageningen University

Claire Vieille, Editor (2020)

Michigan State University

Gerrit Voordouw, Editor (2017)

University of Calgary

Ning-Yi Zhou, Editor (2021)

Shanghai Jiao Tong University

Thomas E. Shenk, Chairman, Journals Board

Stefano Bertuzzi, Chief Executive Officer

Barbara M. Goldman, Director, Journals

Barbara S. Slinker, Production Editor

Stephanie L. D'Agostino, Assistant Production Editor

Michael E. Lerman, Assistant Production Editor

EDITORIAL BOARD

Adly M. M. Abd-Alla (2018)

Lorenz Adrian (2017)

Warish Ahmed (2019)

Birgit E. Alber (2019)

Gladys Alexandre (2018)

Michael S. Allen (2017)

Eric Altermann (2018)

Matthew J. Arduino (2019)

Arthur I. Aronson (2018)

Fred O. Asiegbu (2019)

Shota Atsumi (2018)

Reha O. Azizoglu (2018)

Charles W. Bacon (2018)

Craig Baker-Austin (2018)

Matthew G. Bakker (2019)

David A. Baltrus (2018)

Jeffrey A. Banas (2018)

Ravi D. Barabote (2018)

Jeri D. Barak (2019)

Rodolphe Barrangou (2017)

Josefina Barrera-Cortes (2018)

Mirko Basen (2018)

Dennis A. Bazylinski (2018)

Teresa M. Bergholz (2018)

Stefan Bertilsson (2018)

Jennifer F. Biddle (2018)

Kelly A. Bidle (2019)

Linda F. Bisson (2019)

Christopher Blackwood (2019)

Sara E. Blumer-Schuette (2018)

Paul L. E. Bodelier (2018)

Matthias Boll (2018)

Daniel R. Bond (2018)

Kostas Bourtzis (2018)

E. Fidelma Boyd (2018)

Eric S. Boyd (2018)

Alejandra Bravo (2018)

Frederick Breidt (2017)

Nichole A. Broderick (2018)

Peter A. Bron (2017)

Harald Bruessow (2019)

Harry Brumer (2017)

Andreas Brune (2018)

Alison Buchan (2017)

Wolfgang Buckel (2019)

Daniel H. Buckley (2017)

Bruno Bühler (2019)

William Burkhardt (2018)

Helen Y. Buse (2019)

Stefan G. Buzoianu (2017)

Douglas R. Call (2019)

Amy V. Callaghan (2018)

Barbara J. Campbell (2019)

Isaac K. O. Cann (2018)

Yiping Cao (2017)

Michelle Q. Carter (2018)

Rick Cavicchioli (2019)

Stéphane Chaillou (2018)

Woo-Suk Chang (2018)

John M. Chaston (2018)

Jinru Chen (2017)

Shicheng Chen (2017)

Wilfred Chen (2018)

Yi Chen (2017)

Yuhuan Chen (2018)

Michael L. Chikindas (2019)

Ludmila Chistoserdova (2018)

Konstantin Chumakov (2018)

Kuang-Ren Chung (2017)

Clifford G. Clark (2017)

J. Douglas Cossar (2018)

Donald A. Cowan (2018)

Alison M. Cupples (2019)

Hongyue Dang (2018)

Rolf Daniel (2018)

Michelle Danyluk (2019)

Alexandre J. da Silva (2018)

Jennifer M. DeBruyn (2017)

Svetlana N. Dedysh (2018)

Zixin Deng (2019)

Tom Desmet (2018)

Mickaël Desvaux (2018)

Eduardo Díaz (2019)

Francisco Diez-Gonzalez (2019)

Lubbert Dijkhuizen (2019)

Alan A. DiSpirito (2019)

Jayna L. Ditty (2019)

Jan Dolfing (2018)

James S. G. Dooley (2018)

Angela E. Douglas (2019)

Irina S. Druzhinina (2018)

Doris d'Souza (2017)

Alain C. Dufour (2019)

Anne K. Dunn (2018)

Jonathan Dworkin (2017)

Daniel C. Eastwood (2017)

Leo Eberl (2018)

Elizabeth A. Edwards (2019)

Armin Ehrenreich (2019)

Henry L. Ehrlich (2018)

Patrick Y. Eichenberger (2018)

Mark A. Eiteman (2018)

Kathryn T. Elliott (2018)

Mostafa S. Elshahed (2018)

David Emerson (2018)

Philipp Engel (2019)

Tobias J. Erb (2017)

Danilo Ercolini (2018)

Leonardo Erijman (2019)

Manuel Espinosa-Urgel (2018)

Patrick Fach (2018)

Joseph O. Falkinham (2017)

Séamus Fanning (2018)

Babu Z. Fathepure (2019)

J. Christopher Fenno (2019)

Juan Ferré (2019)

J. Gregory Ferry (2018)

Martin Filion (2018)

Steven E. Finkel (2019)

Steven L. Foley (2017)

Philipp Franken (2018)

Eelco Franz (2019)

W. Florian Fricke (2017)

Michael W. Friedrich (2018)

Barbara E. Funnell (2018)

Vincenzina Fusco (2018)

Shinya Fushinobu (2018)

Cormac G. M. Gahan (2018)

Victor P. J. Gannon (2019)

Michael G. Gänzle (2019)

Jeffrey G. Gardner (2019)

Beilei Ge (2019)

Elke Genersch (2018)

Johannes Gescher (2018)

Lutz Geue (2019)

Lisa M. Gieg (2019)

Jennifer B. Glass (2017)

Emanuel Goldman (2019)

Mark Gomelsky (2017)

Lawrence D. Goodridge (2018)

Lisa A. Gorski (2018)

Yuval Gottlieb (2018)

David E. Graham (2017)

Jeffrey A. Gralnick (2018)

Stefan J. Green (2017)

Leslie A. Gregg-Jolly (2017)

Amy M. Grunden (2018)

Yitzhak Hadar (2019)

Thomas E. Hanson (2018)

Eiji Haramoto (2018)

Steven D. Harris (2017)

Mark E. Hart (2017)

Valerie J. Harwood (2018)

Annele I. Hatakka (2019)

Jianzhong He (2018)

Brian P. Hedlund (2017)

Henry S. Heine (2018)

Thorsten Heinekamp (2018)

Hermann J. Heipieper (2018)

Niels B. Hendriksen (2018)

Kristina L. Hillesland (2018)

Dirk Hoffmeister (2018)

Emily B. Hollister (2018)

Allen L. Honeyman (2018)

Dallas G. Hoover (2018)

Marcus A. Horn (2017)

Adina C. Howe (2018)

John A. Hudson (2017)

Jeroen Hugenholtz (2018)

Hor-Gil Hur (2018)

Michael R. Hyman (2019)

Michael F. Hynes (2019)

Kiyohiko Igarashi (2019)

Michele M. Igo (2018)

Masaharu Ishii (2017)

Masahiro Ito (2017)

Javier A. Izquierdo (2018)

Mohamed A. Jamal (2018)

Edward L. Jarroll (2018)

Dieter Jendrossek (2018)

Michael B. Jenkins (2018)

Francis E. Jenney (2019)

Ian Jenson (2017)

Zhongjun Jia (2017)

Weihong Jiang (2018)

Nianzhi Jiao (2017)

Karyn N. Johnson (2019)

Zackary I. Johnson (2018)

Jessica L. Jones (2018)

Juan L. Jurat-Fuentes (2019)

Klaus Jürgens (2018)

Martin Kaltenpoth (2019)

Yoichi Kamagata (2018)

Boran Kartal (2017)

John J. Kelly (2018)

William R. Kenealy (2018)

Eungbin Kim (2018)

David H. Kingsley (2019)

James E. Kirby (2019)

Martin G. Klotz (2018)

Kalmia E. Kniel (2017)

Ryuichi Koga (2018)

Steffen Kolb (2017)

Konstantinos T. Konstantinidis (2018)

Shige Koseki (2018)

Michael Y. Kosoy (2017)

Balasubrahmanyam Kottapalli (2017)

Sven Krappmann (2018)

Christian P. Kubicek (2018)

Sergei V. Kuchin (2018)

Timothy M. LaPara (2018)

Gisèle LaPointe (2017)

Robert A. LaRossa (2018)

Luis F. Larrondo (2017)

Ray A. Larsen (2018)

Peter C. K. Lau (2018)

Jared R. Leadbetter (2018)

Paulo C. Lemos (2018)

Daniel J. Lessner (2018)

Mark A. Lever (2019)

Jianrong Li (2019)

Wenli Li (2019)

Steven Lindow (2019)

Erin K. Lipp (2018)

Xueduan Liu (2019)

Karen G. Lloyd (2018)

John C. Londesborough (2018)

Alexander Loy (2017)

Xiaonan Lu (2019)

Yahai Lu (2019)

Taina K. Lundell (2018)

Eugene L. Madsen (2018)

Jacques Mahillon (2017)

Mark J. Mandel (2017)

Maria L. Marco (2018)

Julia A. Maresca (2018)

Matthew J. Marshall (2019)

Marta S. L. Martins (2018)

Eiji Masai (2017)

Timothy E. Mattes (2017)

Ann G. Matthysse (2018)

Timothy R. McDermott (2018)

Quinn S. McFrederick (2018)

Michael J. McInerney (2019)

James McKinlay (2018)

Robert J. C. McLean (2018)

Aubrey F. Mendonca (2018)

John S. Meschke (2017)

Chris H. Miller (2018)

Kathryn Milligan-Myhre (2019)

Debra L. Milton (2019)

Kiwamu Minamisawa (2018)

Kentaro Miyazaki (2017)

Robert Moore (2018)

Corrie S. Moreau (2017)

Paula J. Mouser (2018)

Rodney A. Moxley (2019)

Jochen A. Müller (2018)

T. G. Nagaraja (2018)

Yuji Nagata (2018)

Brett A. Neilan (2017)

Josh D. Neufeld (2018)

Irene L. G. Newton (2018)

Nancy N. Nichols (2018)

Jens Nielsen (2018)

Brendan A. Niemira (2018)

Dietrich H. Nies (2018)

Pablo I. Nikel (2019)

Nobuhiko Nomura (2017)

Heléne Norder (2019)

Andrew V. Ogram (2018)

Haley F. Oliver (2019)

Julie B. Olson (2018)

Huub J. M. Op den Camp (2018)

Ivan J. Oresnik (2017)

Kelli L. Palmer (2018)

Konstantinos Papadimitriou (2019)

Hee-Deung Park (2017)

Ilenys M. Perez-Diaz (2018)

Gregg S. Pettis (2017)

Laurent Philippot (2017)

Carmen Pin (2018)

Antonio G. Pisabarro (2017)

Mircea Podar (2017)

Shawn W. Polson (2018)

David L. Popham (2018)

Rolf A. Prade (2018)

M. Auxiliadora Prieto (2018)

Qingsheng Qi (2017)

Ralf Rabus (2017)

Mark Radosevich (2017)

Andreja Rajkovic (2018)

Gemma Reguera (2018)

Bernd H. A. Rehm (2018)

Gregor Reid (2018)

Peter Richard (2018)

Christian Riedel (2018)

Rita V. M. Rio (2018)

Courtney J. Robinson (2018)

Daniel D. Rockey (2019)

Jorge L. M. Rodrigues (2019)

David Rodrígues-Lázaro (2018)

Fernando Rojo (2019)

Margaret F. Romine (2017)

Michael Rother (2018)

Patricia Ruas-Madiedo (2019)

Knut Rudi (2019)

Daâd A. Saffarini (2017)

Irene Salinas (2019)

Irene Sánchez-Andrea (2019)

Robert A. Sanford (2018)

Daisuke Sano (2018)

Anderson S. Sant'Ana (2018)

Brian D. Sauders (2019)

Gary Sawers (2017)

Frank A. Scannapieco (2017)

Birgit E. Scharf (2017)

Herb E. Schellhorn (2019)

Jonathan S. Schilling (2018)

Bernhard Schink (2018)

John W. Schmidt (2019)

Michael G. Schmidt (2018)

Monika Schmoll (2017)

Hildgund Schrempf (2017)

Dirk Schüler (2018)

Bernhard Seiboth (2017)

Yuji Sekiguchi (2018)

David A. Sela (2018)

Jeremy D. Semrau (2017)

John M. Senko (2018)

Peter Setlow (2018)

Luca Settanni (2019)

Robert M. Q. Shanks (2018)

Manan Sharma (2018)

Thomas J. Sharpton (2019)

Cody S. Sheik (2019)

Joshua D. Shrout (2018)

Mark W. Silby (2018)

Mitchell H. Singer (2017)

Steven W. Singer (2018)

David R. Singleton (2018)

Matthias Sipiczki (2019)

Panos N. Skandamis (2019)

Evan W. Skowronski (2018)

J. Leif Smith (2019)

Mario Soberon (2018)

Helena M. Solo-Gabriele (2018)

Greg A. Somerville (2017)

Stephen Spiro (2019)

Lisa Y. Stein (2019)

Alexander Steinbüchel (2017)

Laura Steindler (2019)

Roger Stephan (2017)

Ann M. Stevens (2019)

Brian Stevenson (2019)

George C. Stewart (2018)

Jill R. Stewart (2018)

Michael R. Strand (2019)

Laura K. Strawn (2019)

Wolfgang R. Streit (2018)

Garret Suen (2019)

Matt D. Sweeney (2019)

Teruhiro Takabe (2019)

Ben D. Tall (2017)

Takuji Tanaka (2018)

Ralph S. Tanner (2017)

Gerald W. Tannock (2017)

Sam R. Telford (2018)

Casey M. Theriot (2019)

Michael G. Thomas (2018)

Linda S. Thomashow (2018)

Thomas Thurnheer (2018)

Ming Tien (2018)

Nicolas Toro Garcia (2017)

César I. Torres (2018)

Effie Tsakalidou (2019)

Masataka Tsuda (2018)

Takeharu Tsuge (2018)

Hidetoshi Urakawa (2017)

Wouter R. van der Star (2017)

Jan-Peter van Pijkeren (2018)

Cristian Varela (2018)

Michael L. Vasil (2019)

Marco Ventura (2018)

Karen L. Visick (2019)

Dmitriy V. Volokhov (2018)

Julia A. Vorholt (2018)

Stéphane Vuilleumier (2018)

S. Aljoscha Wahl (2018)

Judy D. Wall (2018)

Gejiao Wang (2018)

Keith Warriner (2017)

J. Scott Weese (2018)

Laure Weisskopf (2018)

Derek H. Wells (2018)

Tingyi Wen (2018)

Volker F. Wendisch (2018)

Steven W. Wilhelm (2018)

Kurt E. Williamson (2017)

David B. Wilson (2019)

Charles F. Wimpee (2017)

Regina-Michaela Wittich (2017)

Gideon M. Wolfaardt (2019)

Randy W. Worobo (2019)

Qinglong Wu (2018)

Chuanwu Xi (2017)

Hua Xiang (2018)

Lihua Xiao (2018)

Luying Xun (2018)

S. Steve Yan (2019)

Liang Yang (2019)

X. Frank Yang (2018)

Yunfeng Yang (2018)

Süleyman Yildirim (2017)

Ken-ichi Yoshida (2018)

Bo Yu (2018)

Daniel R. Zeigler (2018)

Jill H. Zeilstra-Ryalls (2017)

Karsten Zengler (2017)

Qijing Zhang (2017)

Wei Zhang (2018)

Yu-Zhong Zhang (2017)

Jiangchao Zhao (2019)

Ning-Yi Zhou (2018)

Erik R. Zinser (2019)

Erwin G. Zoetendal (2018)

Sergey B. Zotchev (2018)

Peter A. Zuber (2017)

Applied and Environmental Microbiology (ISSN 1098-5336), a publication of the American Society for Microbiology (ASM), 1752 N St., N.W., Washington, DC 20036-2904, is devoted to the advancement and dissemination of applied knowledge as well as ecological knowledge, both applied and fundamental, concerning microorganisms.

Applied and Environmental Microbiology became an online-only journal with volume 82, issue 1 (January 2016). The final print version was volume 81, issue 24 (December 2015). Previous print ISSN: 0099-2240.

Instructions to Authors can be found at http://journalitas.asm.org/t/49536 and are updated throughout the year. Applied and Environmental Microbiology is published twice monthly, one volume per year. For subscription prices, contact the Subscriptions Unit, ASM (asmjournals{at}subscriptionoffice.com). Correspondence relating to reprint orders should be directed to ASM Author Services (ASM.AuthorServicesSupport{at}cenveo.com), and correspondence relating to disposition of submitted manuscripts, proofs, and general editorial matters should be directed to the Journals Department, American Society for Microbiology, 1752 N St., N.W., Washington, DC 20036-2904. Phone: (202) 737-3600 (see http://journals.asm.org/site/misc/staffcontacts.xhtml ). ASM policies for permissions, commercial reprints, and ePrints can be found at http://journals.asm.org/site/misc/reprints.xhtml .

CODEN: AEMIDF

doi:10.1128/AEM.masthead.83-4